Muscle Specific Hypertrophy: Biceps, Back, and Lower Body

In Part 1, we discussed the fiber type composition and biomechanics of the chest, triceps, and shoulders. In this installment, we'll cover the other major muscle groups.

The Biceps & Brachioradialis

The biceps, consisting of a long (outer) and short (inner) head, may seem like a simple muscle with straightforward biomechanics, but it's in fact a tri-articulate muscle that crosses the elbow, forearm, and shoulder.

The short head of the biceps is a shoulder flexor, albeit a weak one, which means it enters active insufficiency when performing elbow flexion and shoulder flexion simultaneously. In other words, it can't perform these two tasks at the same time, so any biceps machine or exercises that has your elbows in front of your shoulders only trains the long head.

The same goes, to a lesser degree, for exercises involving shoulder abduction. So the next time you want to do standing cable curls in front of the mirror in a front double biceps pose, remember that you're likely training your ego more than your biceps. Worry about overall mass first before fussing around with building peaks like Albert Beckles.

Another way to preferentially recruit one head over the other is by manipulating the position of the hands relative to the elbows. A wide grip emphasizes the short head, while a narrow grip emphasizes the long head. A shoulder width grip still emphasizes the long head over the short head.

Again, you should be pretty advanced before even considering this. Most trainees would be best served by sticking with exercises that have their elbows close to their sides.

To emphasize the brachialis over the biceps, perform slow contractions with the biceps in a disadvantageous position (shoulders flexed). The brachialis is more slow-twitch dominant than the biceps, which responds best to fast contractions.

Another consideration for optimal biceps recruitment is hand position. The biceps is very effective with the hands supinated (palms up), somewhat less effective with the hands in a neutral position, and poor with the hands pronated (palms down).

So why are some exercises easier with the hands in neutral position? It's because the brachioradialis, another elbow flexor and one of the larger forearm muscles, is most effective in that position.

Regarding the number of reps to use, while the biceps is fast twitch dominant, it's only dominant by approximately 5%, so medium to low reps work best. The brachioradialis is more fast twitch with ~60% type II fibers, so go heavy on those hammer curls.

Take home message:

  • Stick to standard curls with your elbows at your sides before worrying about emphasizing the brachialis and biceps peak.
  • Use a supinated grip and medium reps for biceps work and a neutral grip and lower reps for the brachioradialis.

The Forearms

The forearms are composed of so many small muscles with varying fiber type compositions and different biomechanics that the full story would go well beyond the scope of this article.

Suffice it to say, elbow extension and flexion cover the largest of the forearm muscles, so do those movements before trying stuff like ulnar deviations and finger exercises.

In accord with their heavy usage, most forearm muscles are slow twitch dominant, so use high reps. Also, be careful of doing more wrist flexion than extension in your programs.

The Lats & Teres Major

The latissimus dorsi and its little helper on top, the teres major, function primarily as shoulder adductors (pulling the elbows into the sides) and shoulder extensors (the movement opposite of a front raise).

Shoulder extension is performed mainly by the higher fibers of the lats, whereas shoulder adduction is performed mainly by the lower fibers of the lats. Maximum lat hypertrophy thus requires the use of both movements.

Many lifters skimp on shoulder adduction, probably because it's harder (as in wide grip pull-ups versus chin-ups), thereby limiting their back width.

A little known fact about the lats is that, just as for the chest, shoulder rotation influences the muscle's biomechanical efficiency. For shoulder adduction, the lats are strongest when the shoulders are somewhat externally rotated. For shoulder extension, the lats are strongest when the shoulders are in neutral position.

If you stick your arms out to the sides, as in the iron cross, you internally rotate your shoulders by pointing your thumbs to the floor and externally rotate them by pointing your thumbs at the ceiling.

Watch the movement of your shoulder. If you use a rope, rings, or any handle that can rotate, you should naturally rotate the shoulder to the optimal position during most pulling exercises. If you don't, you need to use a handle that puts your shoulders in the optimal position yourself. For example, during straight-arm pulldowns, you should use a slightly V-shaped handle instead of a straight bar to maximally recruit your lats.

On the topic of the lats' muscle fiber type composition, the lats on average have an equal amount of slow and fast twitch fibers, so they respond best to medium reps.

Take home message:

  • Include both shoulder extension and adduction in your programs.
  • Think about your grip and shoulder rotation for your exercises.
  • The lats respond best to medium volume.


The Traps

The trapezius is an exemplary muscle. The upper traps are visible in the mirror, but the middle and lower traps aren't, so only the upper traps get attention; a perfect demonstration of the mirror muscle effect.

Even among the people that train their back as hard as their mirror muscles, the traps tend to get left behind. When was the last time you did isolation work for your middle or lower traps?

Although the lower and upper fibers of the trapezius have opposing functions (depress and elevate the scapulae, respectively), training the traps is pretty basic – depress, retract and elevate your scapulae against resistance in equal amounts. (Key phrase being, in equal amounts!)

Also, you may think you're achieving structural balance with your programs because you're pushing and pulling in a 1:1 ratio, but you should really do more pulling volume than pushing.

The chest is fast twitch dominant, but the traps are a postural muscle and are correspondingly slow twitch dominant, so you should do more volume for your traps than for your chest.

It's worth noting that the rhomboids, which also retract the scapulae, are composed of approximately 55% fast twitch fibers, but since these muscles are invisible from the outside, this shouldn't concern bodybuilders.

Also, for those training their necks, the sternocleidomastoid, a neck flexor and extensor, is about 65% fast twitch.

Take home message:

  • Your traps are important. Train them fully, and train them more than your pushing muscles.

The Glutes, Hamstrings, and Lower Back

These muscles comprise the mid-lower posterior chain and have massive force output potential. They often work together during hip extension, where the erector spinae stabilizes the spine, and the glutes and hams move the hips. Besides extending the spine, the erector spinae also laterally flex and rotate the spine.

The gluteus maximus and hams are mainly responsible for extending the hips, but the hamstrings also flex the knee. (The short head of the biceps femoris even performs knee flexion exclusively.) This dual function of the hams has very important implications for the training of your glutes, hams, and calves.

If the biarticulate hamstrings try to stretch at the hip and the knee, they enter passive insufficiency. If they try to flex at the hip and the knee, they enter active insufficiency. Basically, the hams can't perform their functions at both joints at the same time.

This means that during hip extension exercises, such as deadlifts, the more you flex your knees the less the hams can come into play, and the more you emphasize the glutes. If you only do bent-knee exercises for your legs (and no leg curls), you're not completely training your hamstrings.

Then again, many lifters have weak, underactive glutes and strong, overactive hams, so they should do their hip extension movements with bent knees.

Since the glutes and hams both flex the hip, bodybuilders may be inclined to exercise both bodyparts at the same time. This is sub-optimal. The glutes are slow twitch dominant by a few percent, but the hamstrings are composed of a whopping 70% fast twitch fibers.

The erector spinae are slow twitch dominant by around 6%. This means you should use low reps for the hamstrings, but medium to high reps for the lower back and the glutes.

Take home messages:

  • Flexing the knee during hip extension takes the hamstrings out of the movement.
  • Use straight leg movements (or leg curls) for low reps to train the hams and medium to high reps with bent knees to train the glutes and lower back.

The Calves

The long, deeper calf muscle, the soleus, keeps you upright all day and is correspondingly close to 90% slow twitch, so don't bother with reps below 15. The gastrocnemius is often called the fast twitch dominant part of the calves, but that's only compared to the super slow soleus.

The lateral head is indeed fast-twitch dominant, but only by a few percent, and the medial head is actually slow-twitch dominant by a few percent, so use medium volume for the gastrocs. You can emphasize the medial head by pointing your toes out and the lateral head by pointing the toes in.

Both the soleus and the gastrocs perform plantar flexion, but the biarticulate gastrocs also flex the knee, and they can't flex at both joints at the same time. Therefore, you can isolate the hamstrings during knee flexion and the soleus during plantar flexion by plantar flexion and knee flexion, respectively.

The Front Thighs

The front of the thighs consist mostly of the quadriceps. The quads in turn are composed of three main heads – the vastus lateralis, rectus femoris, and the vastus medialis oblique (aka VMO or "teardrop" for you muscleheads).

All heads extend the knee, but the rectus femoris also flexes the hips, so it enters active insufficiency when trying to do both at the same time. To demonstrate this, pull your knee up as high as you can. Now try to extend the knee. Hard, huh? That's active insufficiency of the rectus femoris (and passive insufficiency of the hamstrings).

The VMO and the vastus lateralis also pull the patella in opposite directions during knee extension, so it's important to maintain structural balance between these muscles. There's controversy regarding whether and how preferential recruitment of the VMO over the VL is possible, but healthy trainees should be fine provided they use bilateral and unilateral closed chain exercises with a full range of motion.

Regarding the quads' fiber type composition, it varies significantly per head. The VMO is slightly slow-twitch dominant, the vastus lateralis is approximately 57% fast twitch, and the rectus femoris, the center of the thighs literally and metaphorically, is a 65% fast-twitch powerhouse. Now you know why Olympic weightlifters tend to have large quads. For maximum muscle mass, I recommend mainly low reps but also high variety for the quads.

There's a bunch of other small thigh muscles, including the adductors. I'm not going to cover them all, so put simply, the adductors are about 60% slow twitch and the other hip flexors have a 50-50 fast-slow muscle fiber type composition.

Take home message:

  • For quad training, use a large variation of reps with an emphasis on the lower rep ranges.

The Midsection

The rectus abdominis, aka the six-pack, flexes the spine. The obliques assist the abs in this and laterally flex and rotate the spine. Moreover, along with the transverse abdominis, they create intra-abdominal pressure.

As such, clearly the obliques and transverse abdominis are trained by heavy compound exercises, such as squats and deadlifts. The rectus abdominis, conversely, may remain undertrained without direct work, depending on your exercise selection.

Another consideration is that, anecdotally, the anterior core seems to have limited hypertrophic potential, though its muscles are fast twitch dominant by about 5%. Then there's the debate on whether spinal flexion is healthy, or if extra mass is even aesthetically desirable in the midsection.

With all this in mind, I think beginners should do low rep anti-extension exercises, like planks, rollouts and reverse crunches, to strengthen the anterior core, even if only for the sake of structural balance and learning to properly stabilize the trunk with the core.

After that phase, heavy compound exercises, such as vertical pressing and pulling and anterior load carriage, may be sufficient training for bodybuilders, depending on exercise selection and individual preferences.


And with that, dear friends, you're all set. You're now armed with virtually all the biomechanics and fiber type composition knowledge you'll ever need to optimally recruit your muscles, manipulate exercises to emphasize certain muscles, and select optimal training variable parameters for each muscle.

Now go tweak your training program and get growing!


  1. A biomechanical analysis of anterior load carriage. A M Anderson, K A Meador, L R McClure, D Makrozahopoulos, D J Brooks, G A Mirka. Ergonomics. 2007 Dec;50(12):2104-2117.
  2. Biceps activity during shoulder motion: an electromyographic analysis. K Yamaguchi, K D Riew, L M Galatz, J A Syme, R J Neviaser. Clin Orthop Relat Res. 1997 March; (336): 122–129.
  3. Changes in muscle morphology in chronic trapezius myalgia. R Lindman, M Hagberg, K A Angqvist, K Söderlund, E Hultman, L E Thornell. Scand J Work Environ Health. 1991 October; 17(5): 347–355.
  4. Comparison of different strongman events: trunk muscle activation and lumbar spine motion, load, and stiffness. S M McGill, A McDermott, C M Fenwick. J Strength Cond Res. 2009 Jul;23(4):1148-1161.
  5. Correlation between the proportion of fast fibers in the biceps brachii muscle and the relative torque during elbow fl exion in subjects with clinical hypothesis of myopathy. C G N Back, P C B Oliveira, S M G Mattiello-Rosa , C F R Sobreira, E Z Martinez, A C Mattiello-Sverzut. Rev Bras Fisioter. 2008 March/April; 12(2): 107-112.
  6. Data on the distribution of fibre types in five human limb muscles. An autopsy study. F G Jennekens, B E Tomlinson, J N Walton. J Neurol Sci. 1971 November; 14(3): 245–257.
  7. Data on the distribution of fibre types in thirty-six human muscles. An autopsy study. M A Johnson, J Polgar, D Weightman, D Appleton. J Neurol Sci. 1973 January; 18(1): 111–129.
  8. Effect of isometric upper-extremity exercises on the activation of core stabilizing muscles. Sami P. Tarnanen, Jari J. Ylinen, Kirsti M. Siekkinen, Esko A. Mälkiä, Hannu J. Kautiainen, Arja H. Häkkinen. Arch Phys Med Rehabil. 2008 March; 89(3): 513–521.
  9. Effects of high intensity canoeing training on fibre area and fibre type in the latissimus dorsi muscle. S J Baker, L Hardy. Br J Sports Med. 1989 March; 23(1): 23–26.
  10. Fiber type composition and maximum shortening velocity of muscles crossing the human shoulder. R C Srinivasan, M P Lungren, J E Langenderfer, R E Hughes. Clin Anat. 2007 Mar;20(2):144-9.
  11. Fiber type composition of the vastus lateralis muscle of young men and women. R S Staron, F C Hagerman, R S Hikida, T F Murray, D P Hostler, M T Crill, K E Ragg, K Toma . J Histochem Cytochem. 2000 May; 48(5): 623–629.
  12. Fibre types in human abdominal muscles. T Häggmark, A Thorstensson. Acta Physiol Scand. 1979 Dec;107(4): 319-325.
  13. Fibre types in human lumbar back muscles. A Thorstensson, H Carlson. Acta Physiol Scand. 1987 October; 131(2): 195–202.
  14. Leg extension power and muscle fiber composition. E F Coyle, D L Costill, G R Lesmes. Med Sci Sports. 1979 Spring; 11(1): 12–15.
  15. Muscle fibre type distribution, muscle cross-sectional area and maximal voluntary strength in humans. P Schantz, E Randall-Fox, W Hutchison, A Tydén, P O Astrand. Acta Physiol Scand. 1983 February; 117(2): 219–226.
  16. Muscle fibre type populations of human leg muscles. V. R. Edgerton, J. L. Smith, D. R. Simpson. Histochem J. 1975 May; 7(3): 259–266.
  17. Muscle hypertrophy in bodybuilders. P A Tesch, L Larsson. Eur J Appl Physiol Occup Physiol. 1982; 49(3): 301–306.
  18. Muscle strength from adolescence to adulthood–relationship to muscle fibre types. B Glenmark, G Hedberg, L Kaijser, E Jansson. Eur J Appl Physiol Occup Physiol. 1994; 68(1): 9–19.
  19. Myosin heavy-chain isoform distribution, fibre-type composition and fibre size in skeletal muscle of patients on haemodialysis. S Molsted, I Eidemak, H T Sorensen, J H Kristensen, A Harrison, J L Andersen. Scand J Urol Nephrol. 2007;41(6):539-545.
  20. Rehabilitation of scapular muscle balance: which exercises to prescribe? Ann M. Cools, Vincent Dewitte, Frederick Lanszweert, Dries Notebaert, Arne Roets, Barbara Soetens, Barbara Cagnie, Erik E. Witvrouw. Am J Sports Med. 2007 October; 35(10): 1744–1751.
  21. Strength training and anabolic steroids: a comparative study of the vastus lateralis, a thigh muscle and the trapezius, a shoulder muscle, of strength-trained athletes. A Eriksson. Umeå University Medical Dissertations, Umeå. 2006. Series No 1032.
  22. The influence of variations in muscle fibre composition on muscle strength and cross-sectional area in untrained males. R J Maughan, M A Nimmo. J Physiol. 1984 June; 351: 299–311.
  23. The upper extremity of the professional tennis player: muscle volumes, fiber-type distribution and muscle strength. J Sanchís-Moysi, F Idoate, H Olmedillas, A Guadalupe-Grau, S Alayón, A Carreras, C Dorado, J A L Calbet. Scand J Med Sci Sports. 2010 June; 20(3): 524–534.
  24. Trunk muscle activation during dynamic weight-training exercises and isometric instability activities. N Hamlyn, D G Behm, W B Young. J Strength Cond Res. 2007 Nov;21(4):1108-1112.